Who Defeated The Timurids?

Who Defeated The Timurids? The Aq Qoyunlu conquered most of Iran from the Timurids, and by 1500, the divided and wartorn Timurid Empire had lost control of most of its territory, and in the following years was effectively pushed back on all fronts. Who defeated Taimur? In 1391 Timur pursued Tokhtamysh into the Russian steppes